Asteraceae Checklist

Oregon Vascular Plant Checklist: Asteraceae

by Kenton L. Chambers and Scott Sundberg

May 2000

The Oregon Vascular Plant Checklist, of which this Asteraceae checklist is a part, is intended as a comprehensive reference for the consistent application of botanical names in Oregon. The completed Checklist will provide scientific and common names for all taxa (species, subspecies, and varieties) of native and naturalized vascular plants in the state. Each taxon has been identified as either native or exotic, the latter category including both weeds introduced by human activity and cultivated species that have escaped and are reproducing themselves in the wild. All included taxa have been verified through at least one voucher specimen in the herbaria housed at Oregon State University (in rare cases, published references to specimens in some other herbaria are cited). Because various books may differ in the naming of certain Oregon plants, each accepted name in the Checklist is followed by all the alternative scientific names that are found in ten “standard references” (floras and species lists covering all or part of Oregon). These references are listed below. There are a number of plant taxa that, although reported to occur in Oregon, are not represented by vouchered herbarium specimens at OSU. A list of these is given in Appendix A, under the title Excluded Species. By calling attention to these taxa, and by specifically mentioning when a plant is known from Oregon only by a single herbarium voucher, the authors hope to stimulate other botanists to explore the flora and to report their observations of these and other poorly known native and exotic taxa of vascular plants.

The working Oregon Vascular Plant Checklist is maintained in a Paradox relational database in the Oregon State University Herbarium. The Checklist was originally compiled by Karl Urban with assistance from Kenton Chambers, based mainly on published floras and taxonomic literature available in 1989. More recently, members of the Oregon Vascular Plant Checklist Project (see list of project leaders inside the cover) and invited specialists have been revising this initial list. Over 30 contributors are involved in writing treatments for the updated Checklist. In addition, nine members of the Checklist Advisory Board review taxon treatments as they are received. This Asteraceae checklist was written by Kenton Chambers and Scott Sundberg. The name of the particular author of each generic treatment can be obtained at the OSU Herbarium web site <http://www.orst.edu/dept/botany/herbarium>, or is available upon request.

The Asteraceae checklist will be followed by checklists of other Oregon vascular plant families. When all families have been completed, the lists will be compiled into a single comprehensive volume. We anticipate that the names of some Oregon Asteraceae will be changed for the final volume, and that additional taxa will very likely be included. In the period leading up to publication of the Oregon Vascular Plant Checklist, we will be posting supplemental information on the OSU Herbarium web site.

The Asteraceae is the largest family of vascular plants in Oregon. The checklist includes 574 taxa belonging to 123 genera and 492 species. In addition, 542 synonyms (alternative names) are listed. Over twenty-one percent (124) of the taxa are non-native. Thirty-three taxa are known from only one collection at Oregon State University, which houses the herbaria of this university (OSC), the University of Oregon (ORE), and Willamette University (WILLU). Five taxa are known only from old collections on ship ballast in the Portland area. Genera with the greatest numbers of taxa are Erigeron (48 taxa), Aster (33), Artemisia (31), Senecio (28), Cirsium (20), and Antennaria (19).

Online version of the Checklist (http://www.oregonflora.org).

This online version of the Asteraceae checklist includes additions and changes to the June 1998 first printing. It has been slightly modified from the May 2000 hard copy version. Changes in Appendix B have been incorporated into the checklist itself with the exceptions of two notes on Aster and Senecio. This text version of the checklist will only rarely be updated. The current checklist is in database format and may be accessed in the Oregon Flora Project web site.

Checklist Format

Accepted name: Accepted scientific names are listed in italics and boldface. These consist of Latin names and their botanical authorities. Authority abbreviations follow Authors of Plant Names (Brummitt & Powell 1992).

Synonym(s): A select group of synonyms, in italics, follows the accepted name. These synonyms were treated as accepted names in one or more of the standard references listed below.

Common name(s): One or two English common names are included for each taxon. In general, common names were compiled from the standard references; however, a few have been newly composed for use in this Checklist.

Origin: Plants are considered “native” if they are assumed to have been present in Oregon before the arrival of Euro-American settlers, and are designated as “exotic” if they arrived after that time. Most exotic species are introduced weeds or ornamental plants which have escaped from cultivation.

Comments: In this part we include notes on hybridization and intergradation, morphological variation, quotations from the literature, justification for inclusion on the list (in rare cases where no vouchers were seen), and miscellaneous observations such as localized occurrences and data on single-vouchered taxa.

Standard References for the Oregon Vascular Plant Checklist

FN = Flora of North America (Flora of North America Committee 1993‑present; Asteraceae volume not yet published).

FPN = Flora of the Pacific Northwest (Hitchcock & Cronquist 1973).

GWM = Guide to the plants of the Wallowa Mountains of northeastern Oregon (Mason 1975).

IL = An illustrated flora of the Pacific states (Abrams & Ferris 1940‑1960; Asteraceae: Ferris 1960).

IN = Intermountain flora (Cronquist et al. 1972‑present; Asteraceae: Cronquist 1994).

JPM = The Jepson manual - higher plants of California (Hickman et al. 1993).

KZ = A synonymized checklist of the vascular flora of the United States, Canada, and Greenland (Kartesz 1994).

PEK = A manual of the higher plants of Oregon, 2nd edition (Peck 1961).

PLANTS = The PLANTS database (USDA NRCS 1997).

VP = Vascular plants of the Pacific Northwest (Hitchcock et al. 1955-1969; Asteraceae: Cronquist 1955).

Other Abbreviations

c. = central

Co., cos. = County, counties

coll. = collected, collection

Cr. = Creek

det. = determined or annotated

e. = east

GH = Gray Herbarium (at Harvard University)

Mtn., mtn., mtns. = mountain(s)

n. = north

ORE = University of Oregon Herbarium

orth. = orthographic variant or spelling error

OSC = Oregon State University Herbarium

OSU = herbaria housed at Oregon State University

p., pp. = page(s)

publ. = published, publication

R. = River

s. = south

sp., spp. = species (singular, plural)

ssp., sspp. = subspecies (singular, plural)

US = United States of America or United States National Herbarium

var., vars. = variety, varieties

vs. = versus

w. = west

WILLU = Willamette University Herbarium

State abbreviations are those of the US Postal Service.

Terminology

Ballast: Soil, gravel or water used to stabilize ships when they have little or no cargo. Ballast was dumped in port cities, such as Portland, or the old town Linnton, before cargo was loaded for the return voyage. Plants new to the state sometimes grew from seeds in the ballast. Many “ballast plants” did not become established and may have persisted for only one or a few years.

ex: Used between names of two authorities (e.g. Douglas ex Hook., indicating that David Douglas suggested the name in manuscript and that W. J. Hooker validly published the name).

Implied synonyms at the varietal and subspecific level: In both Peck's Manual (PEK) and Abrams & Ferris’ Illustrated Flora (IL), there are usually no direct citations of the autonym names of "typical" varieties or subspecies within recognized species. Instead, the genus and species binomial is given, followed by a description of the typical variety or subspecies, and additional vars. and sspp. are then named, each with a description that differentiates it from the typical one. We consider the autonym names, or typical names, to be “implied” in these cases.

Illegitimate: Following the name of a synonym, this indicates that the name appears in print but is not in accord with the rules for valid or legitimate publication (see Greuter et al. 1994, International Code of Botanical Nomenclature).

In part in <reference>: Following the name of a synonym, part of the plants referred to this name in the reference are placed in synonymy here, others are accepted in the Checklist under other names.

Misapplied names: Names of species occurring in other regions, which in the cited reference are incorrectly applied to Oregon plants.

Orthographic variant (orth.): Alternative, usually incorrect spelling of a name appearing in a standard reference.

´: Before a species epithet or between two taxon names, indicates sporadic plants of hybrid origin or a series of hybrid populations which are not considered to be stable and self-reproducing.

Bibliography

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Anderson, L.C. 1995. The Chrysothamnus-Ericameria connection (Asteraceae). Great Basin Naturalist 55: 84-88.

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Bayer, R.J. and G.L. Stebbins. 1987. Chromosome numbers, patterns of distribution, and apomixis in Antennaria (Asteraceae: Inuleae). Syst. Bot. 12: 305-319.

Barkley, T.M. 1999. The segregates of Senecio, s.l., and Cacalia, s.l., in the Flora of North America north of Mexico. Sida 18:661-672.

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Cronquist, A. 1994. Intermountain flora. Vascular plants of the Intermountain West, U.S.A., volume 5. Asterales. Bronx: The New York Botanical Garden.

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Hitchcock, C.L. and A. Cronquist. 1973. Flora of the Pacific Northwest. Seattle: University of Washington Press.

Howell, J.T. 1959. Studies in Cirsium - II. Leafl. W. Bot. 9: 9-15.

Jansen, R.K., R.S. Wallace, K.-J. Kim, and K.L. Chambers. 1991. Systematic implications of chloroplast DNA variation in the subtribe Microseridinae (Asteraceae: Lactuceae). Amer. J. Bot. 78: 1015-1027.

Kartesz, J.T. 1994. A synonymized checklist of the vascular flora of the United States, Canada, and Greenland, 2nd edition. Portland: Timber Press.

Mason, G. 1975. Guide to the plants of the Wallowa Mountains of northeastern Oregon. Eugene: Museum of Natural History, University of Oregon.

Nesom, G.L. 1992. Taxonomic notes on Erigeron (Asteraceae: Astereae) of California, Nevada, and Arizona. Phytologia 73: 186-202.

Nesom, G.L. 1994. Review of the taxonomy of Aster sensu lato (Asteraceae: Astereae), emphasizing the New World species. Phytologia 77:141-297.

Nesom, G.L. 1997. Taxonomic adjustments in North American Aster sensu latissimo (Asteraceae) Astereae). Phytologia 82:281-288.

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Ochsmann, J. 1999. What is the American “Centaurea maculosa”? Poster abstract presented at the International Botanical Congress in St. Louis.

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Winward, A.H. and E.D. McArthur. 1995. Lahontan sagebrush (Artemisia arbuscula ssp. longicaulis): a new taxon. Great Basin Naturalist 55: 151-157.

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Acknowledgements

We wish to thank the Oregon State University Department of Botany and Plant Pathology and the Oregon State University Herbarium for providing facilities, and the Native Plant Society of Oregon and numerous individuals and organizations for financial support. We thank John Kartesz for providing us with a copy of his database of Oregon plants, which was compared with our checklist in the final stages of its production. We also thank the many other people who have provided information on, and specimens of, our largest Oregon vascular plant family.

Oregon Vascular Plant Checklist: Asteraceae

by Kenton L. Chambers and Scott Sundberg

Acanthospermum australe (Loefl.) Kuntze

starbur (exotic)

Collected on ballast in "Linton, 1916, J.C. Nelson 975 [GH]" (Blake 1921).

Acanthospermum hispidum DC.

hispid starbur (exotic)

Achillea millefolium L.

Achillea borealis Bong.

Achillea lanulosa Nutt. var. eradiata (Piper) M. Peck

Achillea lanulosa Nutt. var. lanulosa implied in PEK

Achillea millefolium L. var. alpicola (Rydb.) Garrett

Achillea millefolium L. var. californica (Pollard) Jeps.

Achillea millefolium L. var. lanulosa (Nutt.) Piper

Achillea millefolium L. ssp. lanulosa (Nutt.) Piper

Achillea millefolium L. var. litoralis Ehrend. ex Nobs

Achillea millefolium L. var. millefolium

Achillea millefolium L. var. occidentalis DC.

Achillea millefolium L. var. pacifica (Rydb.) G.N. Jones

yarrow, milfoil (both native and exotic populations)

A complex polyploid sp. composed of numerous intergrading ecotypes and ecoclines; the named vars. are too poorly defined to be given taxonomic recognition.

Achyrachaena mollis Schauer

blow wives (native)

Acroptilon repens (L.) DC.

Centaurea repens L.

Russian knapweed, Turkestan thistle (exotic)

Adenocaulon bicolor Hook.

trailplant, pathfinder (native)

Ageratina occidentalis (Hook.) R.M. King & H. Rob.

Eupatorium occidentale Hook.

western boneset (native)

Leaf size and toothing are extremely variable.

Agoseris apargioides (Less.) Greene var. apargioides

Agoseris apargioides (Less.) Greene ssp. apargioides implied in IL

Agoseris hirsuta (Hook.) Greene

woolly agoseris (native)

Type of Leontodon hirsutum Hook. (at Kew) from "sandy banks of the Columbia. Douglas; Scouler" fits this var. according to Cronquist; no recent colls. from OR.

Agoseris apargioides (Less.) Greene var. eastwoodiae (Fedde) Munz

Agoseris apargioides (Less.) Greene ssp. maritima (E. Sheld.) Q. Jones ex Cronquist

Agoseris apargioides (Less.) Greene var. maritima (E. Sheld.) Q. Jones ex Cronquist, illegitimate name

Agoseris maritima E. Sheld.

seaside agoseris (native)

Agoseris aurantiaca (Hook.) Greene

Agoseris aurantiaca (Hook.) Greene var. aurantiaca

orange agoseris, slender agoseris (native)

Agoseris elata (Nutt.) Greene

tall agoseris (native)

Agoseris glauca (Pursh) Raf. var. agrestis (Osterh.) Q. Jones ex Cronquist

field agoseris (native)

Agoseris glauca (Pursh) Raf. var. glauca

pale agoseris, short beaked agoseris (native)

Agoseris glauca (Pursh) Raf. var. laciniata (D.C. Eaton) Smiley

Agoseris glauca (Pursh) Raf. var. parviflora (Nutt.) Rydb.

sagebrush agoseris (native)

Agoseris glauca (Pursh) Raf. var. monticola (Greene) Q. Jones ex Cronquist

Agoseris glauca (Pursh) Raf. var. aspera (Rydb.) Cronquist

Agoseris glauca (Pursh) Raf. var. dasycephala (Torr. & A. Gray) Jeps., misapplied in PEK

mountain agoseris (native)

Agoseris grandiflora (Nutt.) Greene

Agoseris laciniata (Nutt.) Greene

Agoseris plebeja Greene

large flowered agoseris (native)

Agoseris heterophylla (Nutt.) Greene

Agoseris heterophylla (Nutt.) Greene ssp. californica (Nutt.) Piper

Agoseris heterophylla (Nutt.) Greene var. crenulata Jeps.

Agoseris heterophylla (Nutt.) Greene var. cryptopleura Jeps.

Agoseris heterophylla (Nutt.) Greene var. heterophylla

Agoseris heterophylla (Nutt.) Greene ssp. normalis Piper

Agoseris heterophylla (Nutt.) Greene var. turgida (H.M. Hall) Jeps.

annual agoseris (native)

Agoseris retrorsa (Benth.) Greene

spear leaved agoseris (native)

Ambrosia acanthicarpa Hook.

Franseria acanthicarpa (Hook.) Coville

bur ragweed, annual bursage (native)

Ambrosia artemisiifolia L.

Ambrosia artemisiifolia L. var. elatior (L.) Descourt.

annual ragweed, common ragweed (exotic)

Ambrosia chamissonis (Less.) Greene var. bipinnatisecta (Less.) J.T. Howell

Franseria chamissonis Less. var. bipinnatisecta Less.

Franseria chamissonis Less. ssp. bipinnatisecta (Less.) Wiggins & Stockw.

cutleaf beach bur (native)

Ambrosia chamissonis (Less.) Greene var. chamissonis

Franseria chamissonis Less. ssp. chamissonis implied in IL

Franseria chamissonis Less. var. chamissonis implied in PEK

silver beachweed, beach bur (native)

A few sheets in herbaria have plants that are clearly intervarietal hybrids.

Ambrosia psilostachya DC.

western ragweed, perennial ragweed (native)

Ambrosia trifida L.

Ambrosia trifida L. var. trifida

giant ragweed (exotic)

Label on herbarium voucher states that plants are volunteering in waste area near where being grown for pollen.

Anaphalis margaritacea (L.) Benth. & Hook. f.

Anaphalis margaritacea (L.) Benth. & Hook. f. var. occidentalis Greene

Anaphalis margaritacea (L.) Benth. & Hook. f. var. subalpina A. Gray

pearly everlasting (native)

Leaf width, shape, and amount of tomentum on the upper surface vary considerably, but no vars. are worthy of recognition.

Ancistrocarphus filagineus A. Gray

Stylocline filaginea (A. Gray) A. Gray

Stylocline filaginea (A. Gray) A. Gray var. depressa Jeps.

woolly hookfruit (native)

Antennaria anaphaloides Rydb.

tall pussytoes, tall everlasting (native)

Antennaria argentea Benth.

silvery pussytoes, silvery everlasting (native)

Antennaria aromatica Evert

aromatic pussytoes (native)

There is some taxonomic uncertainty about this sp. in OR, as the voucher coll. (Wallowa Co., Marble Mtn., Eagle Cap Wilderness, 1975) has been annotated alternatively as A. aromatica and A. rosea by the monographer, R.J. Bayer.

Antennaria corymbosa E.E. Nelson

meadow pussytoes, flat topped pussytoes (native)

Possibly confused with A. rosea in OR, as there is only the single difference of a dark spot on the involucral bracts of A. corymbosa.

Antennaria dimorpha (Nutt.) Torr. & A. Gray

Antennaria dimorpha (Nutt.) Torr. & A. Gray var. latisquama (Piper) M. Peck, illegitimate name

low pussytoes (native)

Antennaria flagellaris (A. Gray) A. Gray

stoloniferous everlasting, flagellate pussytoes (native)

Antennaria geyeri A. Gray

pinewoods pussytoes, Geyer's pussytoes (native)

Antennaria howellii Greene ssp. howellii

Antennaria neglecta Greene var. howellii (Greene) Cronquist

Howell's pussytoes (native)

Antennaria howellii Greene ssp. neodioica (Greene) R.J. Bayer

Antennaria neglecta Greene var. attenuata (Fernald) Cronquist, in part in FPN

Antennaria neglecta Greene var. neodioica (Greene) Cronquist

field pussytoes (native)

Antennaria howellii Greene ssp. petaloidea (Fernald) R.J. Bayer

Antennaria neglecta Greene var. attenuata (Fernald) Cronquist, in part in FPN

Antennaria pedicellata Greene

Blue Mountains everlasting (native)

Antennaria lanata (Hook.) Greene

Antennaria carpatica (Wahlenb.) Bluff & Fingerh. var. lanata Hook.

woolly pussytoes, woolly everlasting (native)

Antennaria luzuloides Torr. & A. Gray ssp. aberrans (E.E. Nelson) R.J. Bayer & Stebbins

Antennaria luzuloides Torr. & A. Gray var. microcephala (A. Gray) Cronquist

Antennaria microcephala A. Gray

small flowered everlasting (native)

Antennaria luzuloides Torr. & A. Gray ssp. luzuloides

Antennaria luzuloides Torr. & A. Gray var. luzuloides

woodrush pussytoes, silvery brown everlasting (native)

Antennaria media Greene

Antennaria alpina (L.) Gaertn., misapplied in GWM

Antennaria alpina (L.) Gaertn. var. media (Greene) Jeps.

alpine pussytoes (native)

Numerous Cascade Mtns. and Wallowa Mtns. specimens vary between this sp. and A. rosea.

Antennaria racemosa Hook.

raceme pussytoes, slender everlasting (native)

Antennaria rosea Greene

Antennaria microphylla Rydb., misapplied in FPN & IN

Antennaria rosea Greene ssp. confinis (Greene) R.J. Bayer

Antennaria rosea Greene ssp. rosea

rosy pussytoes, rosy everlasting (native)

This apomictic sp. is morphologically highly variable. Two related sexual diploid taxa (A. microphylla Rydb. and A. parvifolia Nutt.) are attributed to OR by Bayer & Stebbins (1987) but have been excluded here due to lack of voucher specimens.

Antennaria stenophylla (A. Gray) A. Gray

narrowleaved pussytoes (native)

Antennaria suffrutescens Greene

shrubby pussytoes, Siskiyou everlasting (native)

Antennaria umbrinella Rydb.

umber pussytoes, brown everlasting (native)

Anthemis arvensis L.

corn chamomile, field chamomile (exotic)

Anthemis cotula L.

mayweed chamomile, dogfennel (exotic)

Anthemis tinctoria L.

yellow chamomile, golden marguerite (exotic)

Arctium lappa L.

great burdock (exotic)

Arctium minus (Hill) Bernh.

common burdock (exotic)

Probably freely hybridizing with A. lappa, according to Cronquist (IN5-386).

Arctium tomentosum Mill.

cotton burdock (exotic)

On basis of key and description in Gleason and Cronquist, Manual of vascular plants of northeastern United States and adjacent Canada, 2nd edition, 1991, pg. 610.

Arnica amplexicaulis Nutt.

Arnica amplexicaulis Nutt. var. amplexicaulis

Arnica amplexicaulis Nutt. var. piperi H. St. John & F.A. Warren

streambank arnica, clasping arnica (native)

The traits of var. piperi are minor differences not worth taxonomic recognition in such a variable genus as Arnica.

Arnica cernua Howell

nodding arnica, serpentine arnica (native)

Sp. is supposed to have glabrous leaves (Wolf & Denford 1984, p. 261), but several colls. having pubescent foliage are present, probably being hybrids with A. spathulata which is sympatric in s.w. OR.

Arnica chamissonis Less. ssp. foliosa (Nutt.) Maguire

Arnica chamissonis Less. var. andina (Nutt.) Ediger & Barkley

Arnica chamissonis Less. var. foliosa (Nutt.) Maguire

narrowleaf arnica, leafy arnica (native)

Arnica chamissonis Less. ssp. incana (A. Gray) Maguire

Arnica chamissonis Less. var. incana (A. Gray) Hultén

silvery arnica (native)

Ssp. incana is a "distinctive ecotype of very wet places" (IN5-094); its velvety pubescence is unique among OR spp. of Arnica.

Arnica cordifolia Hook.

Arnica cordifolia Hook. var. cordifolia

Arnica cordifolia Hook. var. pumila (Rydb.) Maguire

heart leaf arnica (native)

Wolf and Denford (1984, p.270-271) state that var. pumila is an environmentally induced alpine form not worthy of taxonomic recognition.

Arnica discoidea Benth.

Arnica discoidea Benth. var. eradiata (A. Gray) Cronquist

Arnica grayi A. Heller

Arnica parviflora A. Gray ssp. alata (Rydb.) Maguire

Arnica parviflora A. Gray ssp. parviflora implied in PEK

rayless arnica (native)

Wolf & Denford (1984, p. 277) state that A. discoidea should be treated as one highly polymorphic sp. with no infraspecific taxa.

Arnica diversifolia Greene

sticky arnica (native)

Like some other Arnica taxa, this sp. is probably of hybrid origin, one parent being A. mollis, and is maintained by apomictic reproduction; we chose not to use the "´" for this taxon, based on our review of literature.

Arnica fulgens Pursh

hillside arnica, orange arnica (native)

Arnica gracilis Rydb.

Arnica latifolia Bong. var. gracilis (Rydb.) Cronquist

slender arnica (native)

Arnica latifolia Bong.

Arnica latifolia Bong. var. latifolia

broad leaved arnica, mountain arnica (native)

Arnica longifolia D.C. Eaton

Arnica longifolia D.C. Eaton ssp. myriadenia (Piper) Maguire

seep spring arnica, longleaf arnica (native)

Ssp. myriadenia is too poorly defined to be recognized in our treatment for OR.

Arnica mollis Hook.

hairy arnica, cordilleran arnica (native)

In Cascade Mtns. this sp. may be difficult to distinguish from A. amplexicaulis, as only leaf characters separate the two.

Arnica nevadensis A. Gray

Arnica tomentella Greene

Sierra arnica, Nevada arnica (native)

Most herbarium colls. in OR are from the vicinity of Crater Lake; the sp. can be confused with alpine forms of A. cordifolia; for A. tomentella see Wolf & Denford 1984, p. 295.

Arnica parryi A. Gray

Arnica parryi A. Gray var. parryi

Arnica parryi A. Gray ssp. parryi

Arnica parryi A. Gray var. sonnei (Greene) Cronquist

Arnica parryi A. Gray ssp. sonnei (Greene) Maguire

Parry's arnica (native)

Arnica rydbergii Greene

subalpine arnica, Rydberg's arnica (native)

Arnica sororia Greene

twin arnica, bunch arnica (native)

Arnica spathulata Greene

Arnica spathulata Greene ssp. eastwoodiae (Rydb.) Maguire

Arnica spathulata Greene ssp. spathulata implied in IL

spatulate arnica, Klamath arnica (native)

Wolf & Denford (1984, p. 299) view ssp. eastwoodiae as an environmentally induced form of exposed habitats, not worthy of taxonomic recognition.

Arnica viscosa A. Gray

Shasta arnica (native)

Artemisia abrotanum L.

southernwood, garden sagebrush (exotic)

At OSU known only from one OR coll. (Umatilla Co., Pendleton, 1951).

Artemisia absinthium L.

absinthe, wormwood (exotic)

Artemisia arbuscula Nutt. ssp. arbuscula

Artemisia arbuscula Nutt. var. arbuscula

low sagebrush, little sagebrush (native)

Artemisia arbuscula Nutt. ssp. longicaulis Winward & McArthur

Lahontan sagebrush (native)

Based on length of flowering stems, this taxon occurs as far n. as Jefferson Co., OR, although mapped only to extreme s.e. OR by Winward & McArthur (1995).

Artemisia arbuscula Nutt. ssp. longiloba (Osterh.) L.M. Schultz, ined.

Artemisia longiloba (Osterh.) Beetle

early sagebrush (native)

Differs from ssp. arbuscula in its earlier dates of growth initiation and flowering time, blooming in late spring and early summer.

Artemisia arbuscula Nutt. ssp. thermopola Beetle

cleftleaf sagebrush (native)

Winward (1980) cites this taxon from Crook, Wheeler, Grant and Baker cos., whereas L.M. Schultz (JPM-203) does not mention its occurrence in OR.

Artemisia biennis Willd.

biennial wormwood (exotic)

Cronquist (IN5-154) states this sp. is native in n.w. U.S., but JPM-203 cites it as exotic; the type of Willdenow was a garden specimen supposedly from New Zealand, said to be "in error" (G.H. Ward, IL4-408).

Artemisia campestris L. var. scouleriana (Besser) Cronquist

Artemisia campestris L. ssp. pacifica (Nutt.) H.M. Hall & Clem.

Artemisia campestris L. var. purshii (Hook.) Cronquist

Scouler's wormwood, Pacific sagewort (native)

Artemisia campestris L. var. wormskioldii (Besser) Cronquist

Artemisia campestris L. ssp. borealis (Pall.) H.M. Hall & Clem., in part in IL, JPM & KZ

Columbia Islands sagewort (native)

Artemisia cana Pursh ssp. cana

Artemisia cana Pursh ssp. bolanderi (A. Gray) G.H. Ward, misapplied in JPM

silver sagebrush, hoary sagebrush (native)

We agree with Ward (1953) and Cronquist (IN5-164) that the OR material is ssp. cana, not ssp. bolanderi (A. Gray) G.H. Ward as interpreted by Schultz (1986), the latter ssp. being limited to CA.

Artemisia douglasiana Besser

Douglas sagewort, Douglas mugwort (native)

Artemisia dracunculus L.

Artemisia dracunculus L. var. dracunculus

Artemisia dracunculus L. ssp. dracunculus

tarragon, dragon sagewort (native)

Artemisia lindleyana Besser

riverbank wormwood, Columbia River sagewort (native)

Artemisia ludoviciana Nutt. ssp. candicans (Rydb.) D.D. Keck

Artemisia ludoviciana Nutt. var. latiloba Nutt.

gray sagewort (native)

Artemisia ludoviciana Nutt. ssp. estesii K.L. Chambers, in prep.

Estes' artemisia (native)

Artemisia ludoviciana Nutt. ssp. incompta (Nutt.) D.D. Keck

Artemisia ludoviciana Nutt. var. incompta (Nutt.) Cronquist

intermediate sagewort (native)

Artemisia ludoviciana Nutt. ssp. ludoviciana

Artemisia ludoviciana Nutt. var. ludoviciana

Louisiana sagewort, silver wormwood (native)

Artemisia michauxiana Besser

Michaux's mugwort, lemon sagewort (native)

Artemisia nova A. Nelson

Artemisia arbuscula Nutt. var. nova (A. Nelson) Cronquist

Artemisia arbuscula Nutt. ssp. nova (A. Nelson) G.H. Ward

black sagebrush (native)

Artemisia packardiae J.W. Grimes & Ertter

Packard's artemisia, Packard's wormwood (native)

First described in 1979, this sp. is found in our standard references only in IN5-154 and KZ1-069.

Artemisia papposa S.F. Blake & Cronquist

Owyhee sagebrush (native)

At OSU known only from one OR coll. (Malheur Co., Little Owyhee R., 1976).

Artemisia pycnocephala (Less.) DC.

beach wormwood, coastal sagewort (native)

Artemisia rigida (Nutt.) A. Gray

stiff sagebrush, scabland sagebrush (native)

Artemisia spinescens D.C. Eaton

bud sagebrush, spring sagebrush (native)

Artemisia suksdorfii Piper

Suksdorf's sagewort, coastal mugwort (native)

A possible hybrid of this sp. with A. douglasiana was coll. on Sauvies Island by Joseph Howell in 1875 (ORE).

Artemisia tilesii Ledeb. ssp. unalaschcensis (Besser) Hultén

Artemisia hookeriana Besser

Artemisia tilesii Ledeb. var. unalaschcensis Besser

Aleutian mugwort, Cascade wormwood (native)

This taxon is said by Cronquist (VP5-070) to intergrade with A. ludoviciana and A. douglasiana in our region.

Artemisia tridentata Nutt. ssp. tridentata

Artemisia tridentata Nutt. var. tridentata

big sagebrush (native)

Artemisia tridentata Nutt. ssp. vaseyana (Rydb.) Beetle

Artemisia tridentata Nutt. var. vaseyana (Rydb.) B. Boivin

Vasey sagebrush, mountain big sagebrush (native)

Cronquist (FPN-488) had earlier called this the "montane ecotype" of var. tridentata, but in IN5 he raised it to varietal rank.

Artemisia tridentata Nutt. ssp. wyomingensis Beetle & A. Young

Wyoming big sagebrush (native)

Artemisia tripartita Rydb. ssp. tripartita

threetip sagebrush, cutleaf sagebrush (native)

Artemisia vulgaris L.

Artemisia vulgaris L. var. selengensis (Turcz. ex Besser) Maxim.

Artemisia vulgaris L. var. vulgaris

mugwort, lobed wormwood (exotic)

Aster alpigenus (Torr. & A. Gray) A. Gray ssp. alpigenus

Aster alpigenus (Torr. & A. Gray) A. Gray var. alpigenus

Oreostemma alpigenum (Torr. & A. Gray) G.L. Nesom var. alpigenum

alpine aster (native)

Aster alpigenus (Torr. & A. Gray) A. Gray ssp. andersonii (A. Gray) Onno

Aster alpigenus (Torr. & A. Gray) A. Gray var. andersonii (A. Gray) M. Peck

Oreostemma alpigenum (Torr. & A. Gray) G.L. Nesom var. andersonii (A. Gray) G.L. Nesom

Anderson's aster (native)

This ssp. intergrades with ssp. alpigenus in the c. and s. Cascade Mtns. of OR.

Aster alpigenus (Torr. & A. Gray) A. Gray ssp. haydenii (Porter) Cronquist

Aster alpigenus (Torr. & A. Gray) A. Gray var. haydenii (Porter) Cronquist

Oreostemma alpigenum (Torr. & A. Gray) G.L. Nesom var. haydenii (Porter) G.L. Nesom

Hayden's aster (native)

Some plants of this ssp. in the Wallowa Mtns. are indistinguishable from some forms of ssp. alpigenus in the Cascade Mtns.

Aster ascendens Lindl.

Aster adscendens Lindl., orth. in IL

Aster chilensis Nees ssp. adscendens (Lindl.) Cronquist

long leaved aster (native)

Aster breweri (A. Gray) Semple

Chrysopsis breweri A. Gray

Brewer's aster (native)

At OSU known only from one OR coll. (Josephine Co., Bolan Lake, 1945).

Aster brickellioides Greene

Aster brickellioides Greene var. brickellioides

Aster brickellioides Greene var. glabratus Greene

Aster siskiyouensis A. Nelson & J.F. Macbr.

Aster tomentellus (Greene) S.F. Blake ex M. Peck, illegitimate name

Eucephalus brickellioides (Greene) G.L. Nesom

Eucephalus glabratus (Greene) Greene

smooth rayless aster, Siskiyou rayless aster (native)

The amount of leaf pubescence varies even between plants of the the same population, hence var. glabratus is not worth taxonomic recognition.

Aster campestris Nutt.

Aster campestris Nutt. var. bloomeri A. Gray

Aster campestris Nutt. var. campestris

western meadow aster (native)

Aster chilensis Nees

Aster chilensis Nees ssp. chilensis

Aster chilensis Nees var. chilensis

common California aster, Pacific aster (native)

This taxon intergrades with A. subspicatus on the OR coast, but the two spp. usually differ in chromosome number.

Aster conspicuus Lindl.

showy aster (native)

Aster curtus Cronquist

Sericocarpus rigidus Lindl.

rigid white topped aster (native)

Aster cusickii A. Gray

Aster foliaceus Lindl. var. cusickii (A. Gray) Cronquist

Aster foliaceus Lindl. var. lyallii (A. Gray) Cronquist, in part in FPN, IL, PEK & VP

Cusick's aster (native)

Hybrids between A. cusickii and A. occidentalis have been noted in the Wallowa and Blue Mtns. of OR; such plants resemble A. foliaceus var. parryi.

Aster eatonii (A. Gray) Howell

Eaton's aster, Oregon aster (native)

Some high polyploid specimens are suggestive of hybridization with A. subspicatus.

Aster foliaceus Lindl. var. apricus A. Gray

alpine leafybract aster (native)

This var. is a dwarfed alpine form which intergrades with var. parryi.

Aster foliaceus Lindl. var. parryi (D.C. Eaton) A. Gray

Aster foliaceus Lindl. var. canbyi A. Gray, misapplied in FPN, IL, PEK & VP

Aster foliaceus Lindl. var. frondeus A. Gray

Parry's aster, leafybract aster (native)

Aster frondosus (Nutt.) Torr. & A. Gray

Brachyactis frondosa (Nutt.) A. Gray

alkali aster, short rayed aster (native)

Aster gormanii (Piper) S.F. Blake

Eucephalus gormanii Piper

Gorman's aster (native)

Aster hallii A. Gray

Aster chilensis Nees ssp. hallii (A. Gray) Cronquist

Hall's aster (native)

Polyploid intergrades between A. hallii and A. subspicatus occur in the Willamette Valley and Columbia R. Gorge.

Aster hendersonii Fernald

Aster foliaceus Lindl. var. lyallii (A. Gray) Cronquist, in part in JPM

Kootenai aster (native)

Aster hesperius A. Gray

Aster lanceolatus Willd. ssp. hesperius (A. Gray) J. Semple & J. Chmielewski

western willow aster, marsh aster (native)

At OSU known only from one OR coll. (Crook Co., Hwy 126, 10.7 miles w. of Prineville, 1980).

Aster integrifolius Nutt.

thick stemmed aster, entire leaved aster (native)

Aster laevis L.

smooth aster (exotic)

At OSU known only from one OR coll. (Lane Co., s. of Cape Perpetua, 1947, probably escaped from cultivation).

Aster ledophyllus (A. Gray) A. Gray

Aster ledophyllus (A. Gray) A. Gray var. covillei (Greene) Cronquist

Aster ledophyllus (A. Gray) A. Gray var. ledophyllus

Eucephalus ledophyllus (A. Gray) Greene var. covillei (Greene) G.L. Nesom

Eucephalus ledophyllus (A. Gray) Greene var. ledophyllus

Cascade aster (native)

The very minor pubescence difference characterizing var. covillei is not worth taxonomic recognition.

Aster modestus Lindl.

Canadanthus modestus (Lindl.) G.L. Nesom

few flowered aster, great northern aster (native)

Aster nova-angliae L.

New England aster (exotic)

At OSU known only from one OR coll. (Grant Co., Canyon City, 1958).

Aster occidentalis (Nutt.) Torr. & A. Gray

Aster junciformis Rydb., misapplied in PEK

Aster occidentalis (Nutt.) Torr. var. fremontii (Torr. & A. Gray) A.G. Jones

Aster occidentalis (Nutt.) Torr. var. intermedius A. Gray

Aster occidentalis (Nutt.) Torr. var. occidentalis

Aster occidentalis (Nutt.) Torr. var. yosemitanus (A. Gray) Cronquist

Aster paludicola Piper

Aster spathulatus Lindl. var. intermedius (A. Gray) Cronquist

Aster spathulatus Lindl. var. spathulatus, misapplied in IN

western aster, western mountain aster (native)

No distinction seems possible between var. occidentalis and var. intermedius, due to complete intergradation, while var. yosemitanus is an ecological form with linear leaves but no geographical significance, and it is not A. borealis (Torr. & A. Gray) Prov. (A. junciformis).

Aster oregonensis (Nutt.) Cronquist

Aster oregonensis (Nutt.) Cronquist ssp. oregonensis

Sericocarpus oregonensis Nutt.

Sericocarpus oregonensis Nutt. var. oregonensis

Oregon white topped aster (native)

Aster pansus (S.F. Blake) Cronquist

Aster ericoides L. var. pansus (S.F. Blake) B. Boivin

Aster falcatus Lindl. var. crassulus (Rydb.) Cronquist, misapplied in IL

tufted white prairie aster, little gray aster (native)

Aster perelegans A. Nelson & J.F. Macbr.

elegant aster (native)

Aster radulinus A. Gray

rough leaved aster (native)

Aster scopulorum A. Gray

Ionactis alpina (Nutt.) Greene

crag aster, lava aster (native)

Aster sibiricus L. var. meritus (A. Nelson) Raup

Aster meritus A. Nelson

Eurybia merita (A. Nelson) G.L. Nesom

arctic aster, Siberian aster (native)

In OR this taxon is limited to the Wallowa Mtns.

Aster subspicatus Nees

Aster subspicatus Nees var. grayi (Suksd.) Cronquist

Aster subspicatus Nees var. subspicatus

Douglas' aster (native)

Aster vialis (Bradshaw) S.F. Blake

Eucephalus vialis Bradshaw

wayside aster (native)

Baccharis douglasii DC.

saltmarsh baccharis, Douglas’ baccharis (native)

At OSU known only from two OR colls. (Curry Co., lower Chetco R., 1919 and Pistol R., 1999).

Baccharis pilularis DC.

Baccharis pilularis DC. var. consanguinea (DC.) Kuntze

chapparal broom, coyote brush (native)

Balsamorhiza careyana A. Gray var. careyana

Carey's balsamroot (native)

Plants from Hood River and Wasco cos. are placed here rather than in B. deltoidea.

Balsamorhiza careyana A. Gray var. intermedia Cronquist

intermediate balsamroot (native)

Plants of Klamath and Lake cos. are referred to B. deltoidea.

Balsamorhiza deltoidea Nutt.

Puget balsamroot, deltoid balsamroot (native)

Balsamorhiza hookeri Nutt. var. hirsuta (Nutt.) A. Nelson

Balsamorhiza hirsuta Nutt.

hairy balsamroot (native)

Cronquist (IN5-022) no longer maintains B. hirsuta as a distinct sp.; he states "it is the only var. of B. hookeri in n.e. OR."

Balsamorhiza hookeri Nutt. var. hispidula (W.M. Sharp) Cronquist

hispid balsamroot (native)

We follow Cronquist's treatment (IN5) of vars. of B. hookeri; typical var. hookeri is absent from OR, he says (IN5-103).

Balsamorhiza hookeri Nutt. var. idahoensis (W.M. Sharp) Cronquist

Idaho balsamroot (native)

At OSU known only from two OR colls. (Baker Co., Halfway, 1950; and Union Co., Hot Lake, 1923).

Balsamorhiza hookeri Nutt. var. lanata W.M. Sharp

Yreka balsamroot, woolly balsamroot (native)

Reported from Agate Flat, southern Jackson Co., in 1999.

Balsamorhiza hookeri Nutt. var. neglecta (W.M. Sharp) Cronquist

neglected balsamroot (native)

Intergrades with and is difficult to distinguish from B. hookeri var. hispidula.

Balsamorhiza hookeri Nutt. var. platylepis (W.M. Sharp) Cronquist

Balsamorhiza macrolepis W.M. Sharp. var. platylepis (W.M. Sharp) Ferris

Sharp's balsamroot (native)

At OSU known only from one OR coll. (Klamath Co., 8 miles e. of Dairy, 1948).

Balsamorhiza incana Nutt.

Balsamorhiza hookeri Nutt., misapplied in PEK

hoary balsamroot, woolly balsamroot (native)

Balsamorhiza rosea A. Nelson & J.F. Macbr.

rosy balsamroot (native)

Balsamorhiza sagittata (Pursh) Nutt.

arrowleaf balsamroot (native)

This taxon intergrades with B. careyana var. intermedia in c. OR.

Balsamorhiza sericea W.A. Weber

Balsamorhiza macrolepis W.M. Sharp var. platylepis (W.M. Sharp) Ferris, misapplied in IL

Balsamorhiza platylepis W.M. Sharp, misapplied in PEK

silvery balsamroot, silky balsamroot (native)

Hybrids between B. sericea and B. deltoidea occur in Josephine Co., OR.

Balsamorhiza serrata A. Nelson & J.F. Macbr.

serrate balsamroot, toothed balsamroot (native)

A few herbarium sheets are annotated by W.A. Weber as hybrids between this and B. sagittata or B. careyana.

Balsamorhiza ´terebinthacea (Hook.) Nutt.

wormwood balsamroot (native)

We are using this name in a broad sense for hybrids between B. hookeri and any of the three deltoid-leaved spp., B. deltoidea, B. careyana, or B. sagittata. Exact parentage is usually difficult to determine except by field studies.

Balsamorhiza ´tomentosa Rydb.

woolly hybrid balsamroot (native)

This name refers to hybrids between B. incana and B. sagittata (Weber 1953). Ownbey & Weber (1943) discusses two sites in Wallowa Co. where it occurs.

Bellis perennis L.

English daisy, lawn daisy (exotic)

Bidens beckii Torr. ex Spreng.

Megalodonta beckii (Torr. ex Spreng.) Greene

Megalodonta beckii (Torr. ex Spreng.) Greene var. beckii

Megalodonta beckii (Torr. ex Spreng.) Greene var. hendersonii Sherff

Megalodonta beckii (Torr. ex Spreng.) Greene var. oregonensis Sherff

water marigold (exotic)

At OSU known only from two OR colls. (Lane Co., Siltcoos Lake, 1925, 1931; and Klamath Co., Buck Lake, 1897).

Bidens cernua L.

Bidens cernua L. var. cernua

Bidens cernua L. var. elliptica Wiegand

Bidens cernua L. var. minima (Huds.) DC.

nodding beggar ticks, bur marigold (native)

Bidens frondosa L.

leafy beggarticks, sticktight (native)

Sometimes difficult to distinguish from B. vulgata.

Bidens tripartita L.

three lobed beggarticks (exotic)

Herbarium records are late 1800s colls. from Sauvies Island; Multnomah Co., Burlington Bottoms, 1996; and Curry Co., 5 mi. n. of Cape Blanco, 1997.

Bidens vulgata Greene

tall beggarticks, western sticktight (exotic)

Blepharipappus scaber Hook.

Blepharipappus scaber Hook. ssp. laevis (A. Gray) D.D. Keck

Blepharipappus scaber Hook. ssp. scaber

blepharipappus (native)

Ssp. laevis is inconsistent, both morphologically and geographically, and is not worth recognition.

Boltonia asteroides (L.) L'Hér. var. recognita (Fernald & Griscom) Cronquist

boltonia (exotic)

Some colls. with broader phyllaries approach var. latisquama (A. Gray) Cronquist.

Brickellia californica (Torr. & A. Gray) A. Gray

Brickellia californica (Torr. & A. Gray) A. Gray var. californica

California tasselflower (native)

Brickellia grandiflora (Hook.) Nutt.

large flowered tasselflower, large flowered thoroughwort (native)

Brickellia greenei A. Gray

Greene's tasselflower, Greene's thoroughwort (native)

Brickellia microphylla (Nutt.) A. Gray var. microphylla

small leaved brickellia (native)

Brickellia oblongifolia Nutt. var. linifolia (D.C. Eaton) B.L. Rob.

narrowleaf brickellia, narrow leaved thoroughwort (native)

Treatment follows Cronquist (IN5-380). The two vars. intergrade in s.e. OR and few colls. clearly fit in this var.

Brickellia oblongifolia Nutt. var. oblongifolia

narrowleaf brickellia (native)

Cacaliopsis nardosmia (A. Gray) A. Gray ssp. glabrata (Piper) Piper

Luina nardosmia (A. Gray) Cronquist var. glabrata (Piper) Cronquist

tall silvercrown (native)

Cacaliopsis nardosmia (A. Gray) A. Gray ssp. nardosmia

Luina nardosmia (A. Gray) Cronquist

Luina nardosmia (A. Gray) Cronquist var. nardosmia implied in IL

silvercrown (native)

Strother (1978) does not recognize sspp. or vars. of this sp.

Calycadenia fremontii A. Gray

Calycadenia ciliosa Greene

tufted tackweed, Fremont's calycadenia (native)

Not coll. in OR since 1936, per herbarium specimens, but Carr & Carr (1983), sampled one population in Josephine Co., "at jct. of Hwy. 99 and Jump Off Joe Ck. Rds. along Hwy. 5 N."

Calycadenia truncata DC.

Calycadenia truncata DC. var. scabrella (Drew) Jeps.

Calycadenia truncata DC. ssp. scabrella (Drew) D.D. Keck

Calycadenia truncata DC. ssp. truncata

smooth tackweed, rosin weed (native)

Carr & Carr (JPM-219) do not recognize ssp. scabrella as a separate taxon.

Carduus nutans L.

musk thistle (exotic)

Carduus pycnocephalus L.

Italian thistle, compact headed thistle (exotic)

IL4-540 reports the sp. as "local in Lane Co., OR" but no vouchers seen.

Carduus tenuiflorus Curtis

slender flowered thistle (exotic)

Carthamus lanatus L.

Carthamus lanatus L. ssp. lanatus

woolly distaff thistle (exotic)

Carthamus tinctorius L.

safflower (exotic)

Centaurea calcitrapa L.

purple starthistle (exotic)

Centaurea cyanus L.

bachelor's button, cornflower (exotic)

Centaurea diffusa Lam.

diffuse knapweed, tumble knapweed (exotic)

Centaurea iberica Spreng.

Iberian starthistle (exotic)

At OSU known only from one OR coll. (Jackson Co., e. of Medford, 1954).

Centaurea jacea L.

brown knapweed (exotic)

Centaurea melitensis L.

tocalote, Maltese starthistle (exotic)

Centaurea montana L.

montane starthistle (exotic)

Centaurea nigra L.

black knapweed, hardheads (exotic)

Centaurea nigrescens Willd.

Centaurea dubia Suter

short fringed knapweed (exotic)

At OSU known only from one OR coll. (Hood River Co., Hood River, 1932).

Centaurea pratensis Thuill.

meadow knapweed (exotic)

Hybrid sp. derived from C. jacea ´ C. nigra cross.

Centaurea solstitialis L.

yellow star thistle, St. Barnaby's thistle (exotic)

Centaurea stoebe L. ssp. micranthus (Gugler)Hayek

Centaurea biebersteinii DC., possibly misapplied in KZ

Centaurea maculosa, misapplied

spotted knapweed (exotic)

The application of C. biebersteinii DC. to most North American populations of this sp. needs further study.

Name recently changed from C. maculosa based upon research presented at the International Botanical Congress (Ochsmann 1999).

Centaurea virgata Lam.

Centaurea squarrosa Willd.

Centaurea triumfettii All., misapplied in KZ

Centaurea virgata Lam. var. squarrosa (Willd.) Boiss.

squarrose knapweed (exotic)

Treatment follows Wagenitz (1975).

Chaenactis alpina (A. Gray) M.E. Jones

Chaenactis douglasii (Hook.) Hook. & Arn. var. alpina A. Gray

alpine chaenactis, alpine dustymaidens (native)

Chaenactis cusickii A. Gray

Cusick's chaenactis, morning brides (native)

Chaenactis douglasii (Hook.) Hook. & Arn. var. douglasii

Chaenactis douglasii (Hook.) Hook. & Arn. var. achilleaefolia (Hook. & Arn.) A. Nelson, orth. in FPN, IL, IN, PEK & VP

Chaenactis douglasii (Hook.) Hook. & Arn. var. montana M.E. Jones, possibly misapplied in IL

Chaenactis rubricaulis Rydb., possibly misapplied in PEK

dustymaidens, hoary false yarrow (native)

Chaenactis douglasii (Hook.) Hook. & Arn. var. glandulosa Cronquist

sticky headed chaenactis (native)

This var. has distinctive pubescence and is limited to Wallowa Mtns. and adjacent ID plus high Siskiyou Mtns. and adjacent CA.

Chaenactis macrantha D.C. Eaton

large flowered chaenactis, mojave pincushion (native)

Chaenactis nevii A. Gray

John Day chaenactis, Nevius' chaenactis (native)

Chaenactis stevioides Hook. & Arn.

Chaenactis stevioides Hook. & Arn. var. stevioides implied in IL

broad flowered chaenactis, desert pincushion (native)

Chaenactis xantiana A. Gray

desert chaenactis, Xantus' chaenactis (native)

Chaetadelpha wheeleri A. Gray

chaetadelpha, Wheeler's wirelettuce (native)

Chamaemelum mixtum (L.) All.

Anthemis mixta L.

weedy dogfennel (exotic)

Generic assignment as per Flora Europaea 4:165. At OSU known only from one OR coll. (Multnomah Co., Lower Albina, 1902, probably on ballast).

Chondrilla juncea L.

gum succory, rush skeletonweed (exotic)

Chrysanthemum segetum L.

corn marigold, corn chrysanthemum (exotic)

At OSU known only from one OR coll. (Curry Co., Gold Beach, 1928). JPM-228 says it is commonly escaped in coastal urban areas in n.w. CA.

Cichorium intybus L.

wild succory, common chicory (exotic)

Cirsium acanthodontum S.F. Blake

Cirsium remotifolium (Hook.) DC., in part in JPM

Klamath thistle (native)

Two colls. (Douglas and Curry cos.) may be hybrids between this sp. and C. remotifolium.

Cirsium andersonii (A. Gray) Petr.

Anderson's thistle (native)

At OSU known only from one OR coll. (Lake Co., near Roggers Meadow, Warner Mtns., 1991).

Cirsium arvense (L.) Scop.

Cirsium arvense (L.) Scop. var. arvense

Cirsium arvense (L.) Scop. var. horridum Wimm. & Grab.

Cirsium arvense (L.) Scop. var. mite Wimm. & Grab.

Canada thistle, creeping thistle (exotic)

Cronquist (IN5-414) says, "(c)ontemporary European botanists do not consider (the) described variants of the species to be taxonomically significant."

Cirsium brevifolium Nutt.

Palouse thistle (native)

Cirsium brevistylum Cronquist

short styled thistle (native)

Prior to 1953 this sp. went by the name of C. edule Nutt., hence the confusion in some books about the common name, "Indian thistle."

Cirsium callilepis (Greene) Jeps.

Cirsium callilepis (Greene) Jeps. var. callilepis implied in IL

Cirsium callilepis (Greene) Jeps. var. oregonense (Petr.) J.T. Howell

Cirsium callilepis (Greene) Jeps. var. pseudocarlinoides (Petr.) J.T. Howell

Cirsium centaureae (Rydb.) K. Schum.

Cirsium remotifolium (Hook.) DC., in part in JPM

Cirsium remotifolium (Hook.) DC. var. oregonense Petr.

Cirsium remotifolium (Hook.) DC. ssp. pseudocarlinoides Petr.

mountain thistle, fringe bract thistle (native)

Given the considerable infraspecific variation in this sp. and hybridization with other taxa, it is not useful to recognize vars. as done by Howell (1959).

Cirsium canovirens (Rydb.) Petr.

gray green thistle (native)

Cirsium ciliolatum (L.F. Hend.) J.T. Howell

Ashland thistle (native)

Cirsium cymosum (Greene) J.T. Howell

peregrine thistle, Greene's thistle (native)

Cirsium douglasii DC. var. breweri (A. Gray) D.J. Keil & C. Turner

Cirsium breweri (A. Gray) Jeps.

Cirsium douglasii DC. var. canescens (Petr.) J.T. Howell

Brewer's thistle, swamp thistle (native)

At OSU known only from one OR coll. (Lake Co., 15 miles n. of Lakeview, 1927); IN5-394 reports it as "disjunct to a seepy flowing spring area at s.w. base of Steens Mt., in Harney Co., OR."

Cirsium edule Nutt.

Cirsium hallii (A. Gray) M.E. Jones

Indian thistle, edible thistle (native)

The differences between C. edule and C. hallii appear to be minor and inconsistent.

Cirsium occidentale (Nutt.) Jeps. var. candidissimum (Greene) J.F. Macbr.

Cirsium pastoris J.T. Howell

snowy thistle (native)

Cirsium occidentale (Nutt.) Jeps. var. venustum (Greene) Jeps.

Cirsium proteanum J.T. Howell

Venus thistle (native)

At OSU known only from one OR coll. (Josephine Co., bluffs of Illinois R., 15-18 miles below Selma, 1926).

Cirsium peckii L.F. Hend.

Steens Mountain thistle, Peck's thistle (native)

Cirsium pyrenaicum (Jacq.) All.

Pyrenean thistle (exotic)

Identified from description and illustration in Valdés, B., et al., Flora Vascular del Andalucia Occidental, 3:133, 1987; at OSU known only from one OR coll. (Deschutes Co., "Laidlaw" (=Tumalo), 1906).

Cirsium remotifolium (Hook.) DC.

Cirsium remotifolium (Hook.) DC. var. remotifolium

Cirsium remotifolium (Hook.) DC. ssp. remotifolium

few leaved thistle, weak thistle (native)

Two sheets at WILLU are hybrids of this sp. with C. callilepis, from Polk and Yamhill cos.

Cirsium scariosum Nutt.

Cirsium foliosum (Hook.) DC., misapplied in IL, PEK & VP

Cirsium scariosum Nutt. var. scariosum

Cirsium tioganum (Congdon) Petr. var. tioganum

elk thistle (native)

Cirsium subniveum Rydb.

Cirsium utahense Petr., misapplied in FPN, IL, PEK & VP

Jackson Hole thistle, intermountain thistle (native)

The name C. utahense, often used for this sp., is made a synonym of C. neomexicana A. Gray by Cronquist (IN5-398).

Cirsium undulatum (Nutt.) Spreng.

Cirsium undulatum (Nutt.) Spreng. var. undulatum

wavy leaf thistle (native)

Cirsium vulgare (Savi) Ten.

bull thistle, common thistle (exotic)

Cnicus benedictus L.

blessed thistle (exotic)

Columbiadoria hallii (A. Gray) G.L. Nesom

Haplopappus hallii A. Gray

Hall's goldenweed (native)

Conyza bonariensis (L.) Cronquist

Conyza bonariensis (L.) Cronquist var. leiotheca (S.F. Blake) Cuatrec.

Conyza floribunda Kunth

South American conyza (exotic)

Conyza canadensis (L.) Cronquist

Conyza canadensis (L.) Cronquist var. canadensis

Conyza canadensis (L.) Cronquist var. glabrata (A. Gray) Cronquist

Canadian fleabane, horseweed (native)

Degree of stem pubescence is quite variable and does not merit varietal distinction.

Coreopsis lanceolata L.

garden coreopsis (exotic)

At OSU known only from one OR coll. (Curry Co., Brookings, "escaped from gardens," 1929).

Coreopsis tinctoria Nutt. var. atkinsoniana (Douglas ex Lindl.) H.M. Parker

Coreopsis atkinsoniana Douglas ex Lindl.

Columbia coreopsis, calliopsis (native)

Coreopsis tinctoria Nutt. var. tinctoria

golden tickseed, calliopsis (exotic)

E.B. Smith (1975) gives a distribution map showing one site for this var. in central Willamette Valley; presumably escaped from cultivation.

Cotula australis (Sieber ex Spreng.) Hook. f.

Australian cotula (exotic)

Cotula coronopifolia L.

brass buttons (exotic)

Crepis acuminata Nutt.

Crepis acuminata Nutt. ssp. acuminata

long leaved hawksbeard, tapertip hawksbeard (native)

Crepis atribarba A. Heller

Crepis atribarba A. Heller ssp. atribarba

Crepis atribarba A. Heller ssp. originalis Babc. & Stebbins

slender hawksbeard (native)

We agree with Cronquist (IN5-466) that it is not practical to recognize two sspp. of this taxon; the sp. epithet is sometimes misspelled "atrabarba."

Crepis bakeri Greene

Crepis bakeri Greene ssp. bakeri

Crepis bakeri Greene ssp. cusickii (Eastw.) Babc. & Stebbins

Baker's hawksbeard (native)

Crepis barbigera Leiberg ex Coville

bearded hawksbeard (native)

Crepis capillaris (L.) Wallr.

smooth hawksbeard (exotic)

Crepis intermedia A. Gray

gray hawksbeard, intermediate hawksbeard (native)

Crepis modocensis Greene

Crepis modocensis Greene ssp. modocensis

Crepis modocensis Greene ssp. subacaulis (Kellogg) Babc. & Stebbins

low hawksbeard, Modoc hawksbeard (native)

We follow Stebbins (JPM-244) in not attempting to recognize sspp. in the Crepis spp. that are largely apomictic.

Crepis monticola Coville

mountain hawksbeard, Siskiyou hawksbeard (native)

Crepis nana Richardson ssp. ramosa Babc.

Crepis nana Richardson var. ramosa (Babc.) Cronquist

dwarf hawksbeard, tiny hawksbeard (native)

Crepis nicaeensis Balb. ex Pers.

French hawksbeard (exotic)

Crepis occidentalis Nutt.

Crepis occidentalis Nutt. ssp. conjuncta Babc. & Stebbins

Crepis occidentalis Nutt. var. costata A. Gray

Crepis occidentalis Nutt. ssp. costata (A. Gray) Babc. & Stebbins

Crepis occidentalis Nutt. ssp. occidentalis

Crepis occidentalis Nutt. ssp. pumila (Rydb.) Babc. & Stebbins

western hawksbeard (native)

We follow Stebbins (JPM-244) in not recognizing sspp. in Crepis spp. that are largely apomictic.

Crepis pleurocarpa A. Gray

naked stemmed hawksbeard (native)

Crepis pulchra L.

small flowered hawksbeard (exotic)

Crepis runcinata (E. James) Torr. & A. Gray ssp. hispidulosa (Howell ex Rydb.) Babc. & Stebbins

Crepis runcinata (E. James) Torr. & A. Gray var. hispidulosa Howell ex Rydb.

meadow hawksbeard, dandelion hawksbeard (native)

Crepis runcinata (E. James) Torr. & A. Gray ssp. imbricata Babc. & Stebbins

imbricate hawksbeard, Alvord hawksbeard (native)

Crepis setosa Haller f.

rough hawksbeard, bristly hawksbeard (exotic)

Crepis tectorum L.

annual hawksbeard, rooftop hawksbeard (exotic)

Crepis vesicaria L. ssp. taraxacifolia (Thuill.) Thell. ex Schinz & R. Keller

Crepis vesicaria L. ssp. haenseleri (Boiss. ex DC.) P.D. Sell

weedy hawksbeard (exotic)

Crocidium multicaule Hook.

spring gold, gold star (native)

Crupina vulgaris Cass.

crupina, bearded creeper (exotic)

Reported from Umatilla Co. (Couderc-LeVaillant & Roché 1993).

Delairea odorata Lam.

Senecio mikanioides Walp.

German ivy, Cape ivy (exotic)

Established near the coast in Curry Co.; a noxious weed in CA. Use of generic name Delairea is favored by T. Barkley.

Dimeresia howellii A. Gray

dimeresia, doublet (native)

Doronicum plantagineum L.

leopard's bane (exotic)

Escaped from cultivation near Bellfountain, Benton Co.

Eatonella nivea (D.C. Eaton) A. Gray

white eatonella (native)

Erechtites glomerata (Desf. ex Poir.) DC.

Erechtites arguta DC.

New Zealand burnweed, Australian fireweed (exotic)

Erechtites minima (Poir.) DC.

Erechtites prenanthoides (A. Rich.) DC.

Australian burnweed, toothed coast fireweed (exotic)

Ericameria arborescens (A. Gray) Greene

Haplopappus arborescens (A. Gray) H.M. Hall

goldenfleece (native)

Ericameria bloomeri (A. Gray) J.F. Macbr.

Haplopappus bloomeri A. Gray

Haplopappus bloomeri A. Gray var. angustatus A. Gray

Haplopappus bloomeri A. Gray var. bloomeri

Haplopappus bloomeri A. Gray var. sonnei (Greene) H.M. Hall

rabbitbrush goldenweed (native)

This sp. has hairy and glandular forms; it grades into E. greenei.

Ericameria discoidea (Nutt.) G.L. Nesom var. discoidea

Haplopappus macronema A. Gray

Haplopappus macronema A. Gray var. macronema

discoid goldenweed (native)

Ericameria greenei (A. Gray) G.L. Nesom

Haplopappus bloomeri A. Gray var. greenei (A. Gray) Cronquist

Haplopappus greenei A. Gray

Haplopappus greenei A. Gray var. greenei implied in PEK

Haplopappus greenei A. Gray var. mollis A. Gray

Greene's goldenweed (native)

This sp. intergrades with E. bloomeri and may better be considered a var. of it; hairy and glandular forms exist.

Ericameria humilis (Greene) L.C. Anders.

Chrysothamnus humilis Greene

Chrysothamnus viscidiflorus (Hook.) Nutt. ssp. humilis (Greene) C.L. Hitchc. & Cronquist

Chrysothamnus viscidiflorus (Hook.) Nutt. var. humilis (Greene) Jeps.

Truckee green rabbitbrush (native)

Ericameria nana Nutt.

Haplopappus nanus (Nutt.) D.C. Eaton

dwarf heath goldenrod, rubber weed (native)

Ericameria nauseosa (Pall. ex Pursh) G.L. Nesom & G.I. Baird var. hololeuca (A. Gray) G.L. Nesom & G.I. Baird

Chrysothamnus nauseosus (Pall. ex Pursh) Britton ssp. hololeucus (A. Gray) H.M. Hall & Clem.

Chrysothamnus nauseosus (Pall. ex Pursh) Britton var. hololeucus (A. Gray) H.M. Hall

white rabbitbrush (native)

Reports of E. nauseosa ssp. nauseosa (PLANTS) are due to recognition of quadrinomials; var. nauseosa does not occur in OR.

Ericameria nauseosa (Pall. ex Pursh) G.L. Nesom & G.I. Baird var. nana (Cronquist) G.L. Nesom & G.I. Baird

Chrysothamnus nauseosus (Pall. ex Pursh) Britton ssp. nanus (Cronquist) D.D. Keck

Chrysothamnus nauseosus (Pall. ex Pursh) Britton var. nanus Cronquist

little rabbitbrush (native)

Ericameria nauseosa (Pall. ex Pursh) G.L. Nesom & G.I. Baird var. oreophila (A. Nelson) G.L. Nesom & G.I. Baird

Chrysothamnus nauseosus (Pall. ex Pursh) Britton var. artus (A. Nelson) Cronquist

Chrysothamnus nauseosus (Pall. ex Pursh) Britton ssp. consimilis (Greene) H.M. Hall & Clem.

Chrysothamnus nauseosus (Pall. ex Pursh) Britton var. oreophilus (A. Nelson) H.M. Hall & Clem.

Ericameria nauseosa (Pall. ex Pursh) G.L. Nesom & G.I. Baird ssp. consimilis (Greene) G.L. Nesom & G.I. Baird

rubber rabbitbrush (native)

Ericameria nauseosa (Pall. ex Pursh) G.L. Nesom & G.I. Baird var. speciosa (Nutt.) G.L. Nesom & G.I. Baird

Chrysothamnus nauseosus (Pall. ex Pursh) Britton ssp. albicaulis (Nutt.) H.M. Hall & Clem.

Chrysothamnus nauseosus (Pall. ex Pursh) Britton var. albicaulis (Nutt.) Rydb.

Chrysothamnus nauseosus (Pall. ex Pursh) Britton var. speciosus (Nutt.) H.M. Hall

rubber rabbitbrush (native)

Transferred from Chrysothamnus by Nesom & Baird (1993); stabilized hybrids with Ericameria discoidea have been reported from CA (JPM-230).

Ericameria parryi (A. Gray) G.L. Nesom & G.I. Baird var. nevadensis (A. Gray) G.L. Nesom & G.I. Baird

Chrysothamnus parryi (A. Gray) Greene ssp. nevadensis (A. Gray) H.M. Hall & Clem.

Chrysothamnus parryi (A. Gray) Greene var. nevadensis (A. Gray) Hall ex Jeps.

Nevada rabbitbrush (native)

At OSU known only from one OR coll. (Lake Co., s. of Silver Lake Town, 1919).

Ericameria resinosa Nutt.

Haplopappus resinosus (Nutt.) A. Gray

Columbia goldenweed (native)

Ericameria suffruticosa (Nutt.) G.L. Nesom

Haplopappus suffruticosus (Nutt.) A. Gray

shrubby goldenweed (native)

Ericameria viscidiflora (Hook.) L.C. Anders. ssp. lanceolata (Nutt.) L.C. Anders.

Chrysothamnus viscidiflorus (Hook.) Nutt. ssp. lanceolatus (Nutt.) H.M. Hall & Clem.

Chrysothamnus viscidiflorus (Hook.) Nutt. var. lanceolatus (Nutt.) Greene

lanceleaf green rabbitbrush, green rabbitbrush (native)

Transferred from Chrysothamnus by Anderson (1995).

Ericameria viscidiflora (Hook.) L.C. Anders. ssp. puberula (D.C. Eaton) L.C. Anders.

Chrysothamnus viscidiflorus (Hook.) Nutt. ssp. puberulus (D.C. Eaton) H.M. Hall & Clem.

Chrysothamnus viscidiflorus (Hook.) Nutt. var. puberulus (D.C. Eaton) Jeps.

downy green rabbitbrush (native)

Ericameria viscidiflora (Hook.) L.C. Anders. ssp. viscidiflora

Chrysothamnus viscidiflorus (Hook.) Nutt. var. pumilus (Nutt.) Jeps.

Chrysothamnus viscidiflorus (Hook.) Nutt. ssp. pumilus (Nutt.) H.M. Hall & Clem.

Chrysothamnus viscidiflorus (Hook.) Nutt. var. stenophyllus (A. Gray) H.M. Hall

Chrysothamnus viscidiflorus (Hook.) Nutt. ssp. stenophyllus (A. Gray) H.M. Hall & Clem.

Chrysothamnus viscidiflorus (Hook.) Nutt. ssp. viscidiflorus

Chrysothamnus viscidiflorus (Hook.) Nutt. var. viscidiflorus

Ericameria viscidiflora (Hook.) L.C. Anders. var. stenophylla (A. Gray) L.C. Anders.

yellow rabbitbrush, sticky flowered rabbitbrush (native)

Erigeron acris L. var. debilis A. Gray

Trimorpha acris (L.) A. Gray var. debilis (A. Gray) G.L. Nesom

northern daisy (native)

Erigeron acris L. var. kamtschaticus (DC.) Herder

Erigeron acris L. var. asteroides (Andrz. ex Besser) DC., misapplied in FPN, IL, PEK & VP

Trimorpha acris (L.) G.L. Nesom var. kamtschatica (DC.) G.L. Nesom

bitter fleabane (native)

Erigeron aliceae Howell

Eastwood's daisy, Alice's fleabane (native)

Erigeron annuus (L.) Pers.

annual fleabane, sweet scabrous erigeron (exotic)

Erigeron aphanactis (A. Gray) Greene var. aphanactis

rayless shaggy erigeron, basin rayless daisy (native)

Erigeron bloomeri A. Gray var. bloomeri

scabland fleabane, Bloomer's daisy (native)

Erigeron bloomeri A. Gray var. nudatus (A. Gray) Cronquist

Waldo daisy (native)

This is a very weak var.; plants from type locality may have pubescent involucres, and glabrous and strigose leaves may arise from the same root.

Erigeron breweri A. Gray var. klamathensis G.L. Nesom

Klamath daisy (native)

This taxon differs from E. foliosus var. confinis only in its spreading stem pubescence; some intermediates occur in s. Cascade Mtns. of OR.

Erigeron cascadensis A. Heller

Cascade daisy, horseweed (native)

This sp. is closely related to E. leibergii Piper, of WA Cascade Mtns.; the pubescence differences are inconsistent.

Erigeron cervinus Greene

Erigeron delicatus Cronquist

deer fleabane, Siskiyou daisy (native)

Erigeron chrysopsidis A. Gray var. austiniae (Greene) G.L. Nesom

Erigeron austiniae Greene

Erigeron chrysopsidis A. Gray ssp. austiniae (Greene) Cronquist

Austin's fleabane (native)

Erigeron chrysopsidis A. Gray var. brevifolius Piper

Erigeron chrysopsidis A. Gray ssp. brevifolius Piper

alpine yellow daisy, Wallowa daisy (native)

Erigeron chrysopsidis A. Gray var. chrysopsidis

Erigeron chrysopsidis A. Gray ssp. chrysopsidis

golden fleabane, golden daisy (native)

Erigeron compositus Pursh var. compositus

cutleaf daisy, fernleaf fleabane (native)

This var. is rare in our herbaria, unless some high elevation linear-leaflet types are included in it (e.g., Jackson Co., Mt. Ashland).

Erigeron compositus Pursh var. discoideus A. Gray

trifid mountain fleabane (native)

This taxon may be only a growth form of E. vagus.

Erigeron compositus Pursh var. glabratus Macoun

dwarf mountain fleabane (native)

Erigeron corymbosus Nutt.

foothill daisy, longleaf fleabane (native)

Erigeron coulteri Porter

Coulter's daisy, Coulter's fleabane (native)

Erigeron decumbens Nutt. var. decumbens

Pacific fleabane, Willamette Valley daisy (native)

Erigeron disparipilus Cronquist

Snake River daisy (native)

Differs from E. chrysopsidis mainly in white, not yellow, ray florets; at OSU known only from one OR coll. (Wallowa Co., Fairchild Point area, 1991).

Erigeron divergens Torr. & A. Gray var. divergens

diffuse daisy, spreading fleabane (native)

Erigeron eatonii A. Gray var. lavandulus Strother & Ferlatte

Eaton's lavender daisy (native)

Specimens from Clackamas and Marion cos. provisionally referred here by Strother & Ferlatte (1988) may be a montane race of E. decumbens.

Erigeron eatonii A. Gray var. plantagineus (Greene) Cronquist

Erigeron decumbens Nutt. ssp. robustior Cronquist, misapplied in PEK

plantain leaved daisy (native)

Some OR specimens of this were earlier misnamed E. decumbens ssp. robustior by Cronquist.

Erigeron eatonii A. Gray var. villosus (Cronquist) Cronquist

Eaton's shaggy fleabane, Eaton's shaggy daisy (native)

Erigeron elegantulus Greene

volcanic daisy, blue dwarf fleabane (native)

Erigeron engelmannii A. Nelson var. davisii (Cronquist) Cronquist

Erigeron engelmannii A. Nelson ssp. davisii Cronquist

Engelmann's daisy (native)

Erigeron filifolius (Hook.) Nutt. var. filifolius

threadleaf fleabane (native)

Erigeron filifolius (Hook.) Nutt. var. robustior M. Peck

Peck's threadleaf fleabane (native)

Erigeron foliosus Nutt. var. confinis (Howell) Jeps.

Erigeron foliosus Nutt. var. hartwegii (Greene) Jeps., misapplied in FPN, IL, PEK & VP

leafy fleabane (native)

OR plants formerly called var. hartwegii are only a robust growth-form of var. confinis.

Erigeron glaucus Ker Gawl.

beach fleabane, seaside daisy (native)

Erigeron howellii (A. Gray) A. Gray

Howell's daisy, Howell's fleabane (native)

Erigeron inornatus (A. Gray) A. Gray var. inornatus

California rayless daisy, unadorned fleabane (native)

Erigeron linearis (Hook.) Piper

desert yellow daisy, lineleaf fleabane (native)

Collectors' comments on herbarium sheets suggest that this sp. may hybridize with E. bloomeri; some sheets are a mixture of the two spp.

Erigeron lonchophyllus Hook.

Trimorpha lonchophylla (Hook.) G.L. Nesom

spear leaved fleabane, short rayed daisy (native)

Erigeron oreganus A. Gray

gorge daisy, Oregon fleabane (native)

Erigeron peregrinus (Pursh) Greene var. callianthemus (Greene) Cronquist

Erigeron peregrinus (Pursh) Greene var. angustifolius (A. Gray) Cronquist

Erigeron peregrinus (Pursh) Greene ssp. callianthemus (Greene) Cronquist

Erigeron peregrinus (Pursh) Greene var. eucallianthemus Cronquist

peregrine fleabane (native)

Erigeron peregrinus (Pursh) Greene var. peregrinus

Erigeron peregrinus (Pursh) Greene ssp. peregrinus

wandering daisy (native)

In OR this var. occurs only in the higher elevations of the North Coast Range.

Erigeron peregrinus (Pursh) Greene var. scaposus (Torr. & A. Gray) Cronquist

Erigeron ursinus D.C. Eaton, misapplied in GWM

subalpine fleabane (native)

This is the dwarf ecotype of the sp.; Nesom (1992) feels that neither var. angustifolius nor var. scaposus is worth taxonomic recognition.

Erigeron petrophilus Greene var. petrophilus

Erigeron petrophilus Greene var. viscidulus (A. Gray) G.L. Nesom, misapplied in JPM

cliff fleabane, rock daisy (native)

Erigeron philadelphicus L.

Erigeron philadelphicus L. var. philadelphicus

Philadelphia fleabane, Philadelphia daisy (native)

Erigeron poliospermus A. Gray var. poliospermus

cushion fleabane, hairy seeded daisy (native)

Erigeron pumilus Nutt. var. intermedius Cronquist

Erigeron pumilus Nutt. var. euintermedius Cronquist

Erigeron pumilus Nutt. var. gracilior Cronquist

Erigeron pumilus Nutt. ssp. intermedius Cronquist

shaggy fleabane (native)

Vars. intermedius and gracilior intergrade completely; it is useless to try to distinguish them in OR.

Erigeron simplex Greene

alpine daisy, one stemmed fleabane (native)

Known in OR only in high Wallowa Mtns.

Erigeron speciosus (Lindl.) DC. var. speciosus

Erigeron speciosus (Lindl.) DC. var. macranthus (Nutt.) Cronquist, misapplied in GWM & PEK

Erigeron subtrinervis Rydb., misapplied in GWM

showy fleabane, showy daisy (native)

Cronquist annotated some Wallowa Mtns. colls. as "passing to var. macranthus," but that var. itself is not in OR.

Erigeron strigosus Muhl. ex Willd. var. septentrionalis (Fernald & Wiegand) Fernald

prairie fleabane (exotic)

Cronquist (VP5-193) believes that plants referable to this var. in far w. US. are independent hybrids of E. annuus ´ E. strigosus; var. septentrionalis is native in e. US.

Erigeron strigosus Muhl. ex Willd. var. strigosus

daisy fleabane, branching daisy (exotic)

Erigeron tener (A. Gray) A. Gray

slender daisy, delicate erigeron (native)

Very rarely coll.; known in OR only from Steens Mtn. and Snake R., Baker Co.

Erigeron vagus Payson

loose daisy, rambling fleabane (native)

This grows sympatrically with the related sp. E. compositus in high Wallowa Mtns.

Eriophyllum lanatum (Pursh) J. Forbes var. achillaeoides (DC.) Jeps.

yarrow leaved eriophyllum (native)

Eriophyllum lanatum var. grandiflorum (A. Gray) Jeps.

largeflower eriophyllum (native).

At OSU known only from one OR coll. (Curry Co., Pollywog Butte, 1932); intergrades with var. achillaeoides in s. Josephine Co.

Eriophyllum lanatum (Pursh) J. Forbes var. integrifolium (Hook.) Smiley

Oregon sunshine (native)

Intergrades with var. achillaeoides in Klamath & Lake cos. and with var. lanatum in Blue Mtns.

Eriophyllum lanatum (Pursh) J. Forbes var. lanatum

Eriophyllum lanatum (Pursh) J. Forbes var. leucophyllum (DC.) W.R. Carter

common eriophyllum, common woolly sunflower (native)

Eriophyllum lanatum (Pursh) J. Forbes var. lanceolatum (Howell) Jeps.

lanceleaf eriophyllum (native)

Intergrades with var. achillaeoides in Josephine and Curry cos.

Eriophyllum staechadifolium Lag.

Eriophyllum stoechadifolium Lag., orth. in KZ

seaside woolly sunflower, lizardtail eriophyllum (native)

This sp. has seldom been collected in OR in recent years; perhaps overlooked.

Euthamia occidentalis Nutt.

Solidago occidentalis (Nutt.) Torr. & A. Gray

western goldenrod (native)

Filago arvensis L.

field filago (exotic)

At OSU known only from two OR colls. (Wallowa Co., Lostine River Rd., 1970; Morrow Co., Ione area, 1998).

Filago gallica L.

narrowleaf filago, narrow leaved cudweed (exotic)

In OR known only from Curry Co.

Filago minima (Sm.) Pers.

small cudweed (exotic)

At OSU known from only one OR coll. (Lane Co., Blue River Reservoir, 1971, det. by J. Morefield).

Filago vulgaris Lam.

Filago germanica L., non Huds.

herba impia, common cudweed (exotic)

At OSU known only from one OR coll. (Douglas Co., Roseburg, 1937, det. by J. Morefield, who says it is the first record for the Pacific states).

Gaillardia aristata Pursh

blanket flower, great flowered gaillardia (native)

Galinsoga parviflora Cav.

Galinsoga parviflora Cav. var. parviflora

small flowered galinsoga (exotic)

Canne (1977) cites a Brandegee coll. (GH) from Portland.

Galinsoga quadriradiata Ruiz & Pav.

Galinsoga ciliata (Raf.) S.F. Blake

quickweed, ciliate galinsoga (exotic)

Glyptopleura marginata D.C. Eaton var. marginata

carved seed (native)

Gnaphalium californicum DC.

Pseudognaphalium californicum (DC.) Anderb.

California cudweed, California everlasting (native)

Mostly coastal, n. to Clatsop Co., but also in Willamette Valley.

Gnaphalium canescens DC. ssp. thermale (E.E. Nelson) Stebbins & D.J. Keil

Gnaphalium microcephalum Nutt. var. thermale (E.E. Nelson) Cronquist

slender cudweed (native)

Gnaphalium collinum Labill.

Euchiton gymnocephalus (DC.) Anderb.

creeping cudweed (exotic)

In OR known only from coastal Curry Co.

Gnaphalium japonicum Thunb.

Euchiton japonicus (Thinb.) Anderb.

Japanese cudweed (exotic)

Gnaphalium luteoalbum L.

Pseudognaphalium luteoalbum (L.) Hilliard & B.L. Burtt

weedy cudweed (exotic)

At OSU known only from one OR coll. (Multnomah Co., Lower Albina, Portland, 1902). Hyphen removed from sp. epithet as per ICBN, Art. 60.9 (Greuter et al. 1994).

Gnaphalium macounii Greene

Gnaphalium viscosum Kunth, misapplied in FPN & VP

Pseudognaphalium viscosum (Kunth) W.A. Weber, misapplied in PLANTS

sticky cudweed, winged cudweed (native)

Gnaphalium palustre Nutt.

lowland cudweed (native)

Gnaphalium purpureum L.

Gamochaeta purpurea (L.) Cabrera

Gamochaeta ustulata (Nutt.) G.L. Nesom

Gnaphalium purpureum L. var. purpureum

purple cudweed (native)

Gnaphalium stramineum Kunth

Gnaphalium chilense Spreng.

Pseudognaphalium stramineum (Kunth) W.A. Weber

cotton batting cudweed, cotton batting plant (native)

Gnaphalium uliginosum L.

marsh cudweed (exotic)

Grindelia hirsutula Hook. & Arn. var. davyi (Jeps.) M.A. Lane

Grindelia camporum Greene, misapplied in IL

Davy's gumplant (native)

We follow annotations by M.A. Lane and her treatment for JPM in placing this taxon in Josephine and Jackson cos.; it may be difficult to distinguish from G. nana.

Grindelia integrifolia DC.

Grindelia integrifolia DC. var. integrifolia

Grindelia nana Nutt. var. integrifolia Nutt., misapplied in PEK

Willamette Valley gumweed, Puget Sound gumplant (native)

Grindelia integrifolia ´ Grindelia nana var. nana

Willamette hybrid gumplant (native)

Includes many colls. in the Willamette Valley that are morphologically intermediate between the parent spp.

Grindelia nana Nutt. var. discoidea (Nutt.) A. Gray

Grindelia columbiana (Piper) Rydb.

Grindelia nana Nutt. ssp. columbiana Piper

Columbia gumplant, Columbia gumweed (native)

Grindelia nana Nutt. var. nana

Grindelia nana Nutt. var. integerrima (Rydb.) Steyerm.

Grindelia nana Nutt. var. integrifolia Nutt.

Grindelia nana Nutt. ssp. nana implied in IL

Idaho gumweed, low gumweed (native)

Grindelia squarrosa (Pursh) Dunal var. serrulata (Rydb.) Steyerm.

curlycup gumweed, resinweed (native)

Some authors (e.g. Keck in IL4-258, Lane in JPM-274) suggest that this taxon is native farther e. and only introduced in the Pacific states; Cronquist (IN5-256) says it is "characteristic of the Great Basin."

Grindelia stricta DC. var. stricta

Grindelia arenicola Steyerm.

Grindelia integrifolia DC. var. macrophylla (Greene) Cronquist

Grindelia stricta DC ssp. stricta implied in IL

coastal gumweed, Oregon gumplant (native)

Gutierrezia sarothrae (Pursh) Britton & Rusby

broom snakeweed, matchweed (native)

Hazardia whitneyi (A. Gray) Greene var. discoidea (J.T. Howell) W.D. Clark

Haplopappus whitneyi A. Gray

Haplopappus whitneyi A. Gray ssp. discoideus (J.T. Howell) D.D. Keck

Whitney's goldenweed (native)

Helenium autumnale L. var. grandiflorum (Nutt.) Torr. & A. Gray

large flowered sneezeweed, common sneezeweed (native)

Helenium autumnale L. var. montanum (Nutt.) Fernald

Helenium autumnale L. var. autumnale implied in PEK

mountain sneezeweed (native)

Helenium bigelovii A. Gray

tall sneezeweed, Bigelow's sneezeweed (native)

Helenium bolanderi A. Gray

coast sneezeweed (native)

The only differences from H. bigelovii are the slightly longer pappus and the coastal distribution of H. bolanderi.

Helenium puberulum DC.

rosilla (exotic)

Bierner (1972 and in litt.) cites one Willamette Valley coll. for this CA taxon [Haskin 4 (US), Linn Co., Brownsville, 1923].

Helianthella californica A. Gray var. nevadensis (Greene) Jeps.

California false sunflower, California helianthella (native)

At OSU known only from one OR coll. (Douglas Co., head of Elk Cr., 1899).

Helianthella quinquenervis (Hook.) A. Gray

nodding helianthella, fivenerved helianthella (native)

At OSU known only from one OR coll. (Lake Co., Warner Range, 1896).

Helianthella uniflora (Nutt.) Torr. & A. Gray var. douglasii (Torr. & A. Gray) W.A. Weber

Douglas helianthella, false sunflower (native)

Helianthella uniflora (Nutt.) Torr. & A. Gray var. uniflora

Rocky Mountain helianthella, common little sunflower (native)

Rare in OR. Weber (1952) cites two colls. from Harney Co.

Helianthus annuus L.

Helianthus annuus L. ssp. annuus implied in IL

Helianthus annuus L. ssp. lenticularis (Douglas ex Lindl.) Cockerell

common sunflower (native)

We follow JPM and IN5 in not recognizing sspp. or vars. in H. annuus (versus IL4-113). According to Heiser (1969) the distribution of annual sunflower spp. has been strongly affected by humans, hence, the native ranges are unknown.

Helianthus bolanderi A. Gray

Bolander's sunflower (native)

This sp. in s.w. OR is not limited to serpentine, as the "exilis" form is in CA (Heiser, 1949).

Helianthus cusickii A. Gray

Cusick's sunflower, turniproot sunflower (native)

Specimens of Helianthella uniflora may be misidentified as this sp.

Helianthus nuttallii Torr. & A. Gray ssp. nuttallii

Helianthus nuttallii Torr. & A. Gray var. nuttallii

Nuttall's sunflower, cordilleran sunflower (native)

Helianthus petiolaris Nutt.

Helianthus petiolaris Nutt. var. petiolaris implied in IL

Helianthus petiolaris Nutt. ssp. petiolaris

prairie sunflower (exotic)

All references state that this sp. is native farther e. than OR but adventive here. At OSU known only from one OR coll. (Lake Co., three miles s. of Silver Lake, 1963). We follow Cronquist (IN5-30) in not formally accepting sspp. or vars. of this sp.

Helianthus tuberosus L.

Jerusalem artichoke (exotic)

At OSU known only from one OR coll. (Benton Co., Corvallis, 1955, "disturbed area near trash heap, possibly escaped from cultivation, perhaps cultivated intentionally.").

Hemizonia congesta DC. ssp. clevelandii (Greene) Babc. & H.M. Hall

Hemizonia clevelandii Greene

Cleveland's hemizonia, Cleveland's tarweed (native)

All herbarium sheets at OSU are from 1930s or earlier, except one (Josephine Co., road to Babyfoot Lake, 1977).

Hemizonia fitchii A. Gray

Fitch's spikeweed (native)

Hemizonia pungens (Hook. & Arn.) Torr. & A. Gray ssp. pungens

Hemizonia pungens (Hook. & Arn.) Torr. & A. Gray var. pungens

common spikeweed (exotic)

Hemizonia pungens (Hook. & Arn.) Torr. & A. Gray ssp. septentrionalis D.D. Keck

Hemizonia pungens (Hook. & Arn.) Torr. & A. Gray var. septentrionalis (D.D. Keck) Cronquist

western spikeweed (exotic)

Hesperevax acaulis (Kellogg) Greene var. acaulis